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http://palaeo-electronica.org/content/fc-8 Kohli et al., 2016
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The attribution to the Rhaeto-Liassic is established by the stratigraphic position of the Bayreuth Formation between well-dated Keuper sediments and the Gryphaeensandstein Formation (Bloos et al., 2005), and by plant index fossils (Weber, 1968).
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|primary fossil used to date this node|
The wing venation of this fossil is nearly identical to other fossils of the genus Liassophlebia and shares the following synapomorphies with Heterophlebioptera and Heterophlebioidea (Nel et al., 1993; Bechly, 1996): unique unicellular anal loop, which lies beneath the subdiscoidal cell and is ventrally closed by CuAb that is parallel to AA and thus directed towards the wing base instead of the posterior wing margin; subdiscoidal cell of the hind wing with a convex curved or angulated posterior margin; antenodal crossveins between costal margin and ScP suppressed distal of ax2; IR2 arising on RP3/4. The attribution to the stem group of Anisoptera is well established by the unique synapomorphy of the shape of the hind wing discoidal cell, which foreshadows the anisopterid division into hypertriangle and triangle (only sometimes lacking the dividing crossvein).
Kohli, M.K., Ware, J.L., and Bechly, G. 2016. How to date a dragonfly: Fossil calibrations for odonates. Palaeontologia Electronica, 19.1.1FC.
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